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Graduate Biomedical Sciences Programs

Graduate Programs


The mission of the Department of Biomedical Sciences graduate programs is to provide excellent academic programs that educate and train the next generation of scientists, educators, and health-related professionals through scholarship and research that fosters creativity, discovery, and community service.


The programs are flexible and cater to the needs and interests of individual students. Education is predominant through participation in research, taking advantage of opportunities for strategic collaboration between the basic and clinical sciences.

Research Areas in the Biomedical Sciences Department

The research of the faculty covers a wide range of disciples ranging from biochemistry, molecular and cellular biology, organismal biology and translational research. Broadly the department is focused on two major thematic areas: neuroscience of hearing and hearing loss and cancer biology. The Department encourages collaborative research interaction with faculty in the Departments of Pharmacology, Medical Microbiology & Immunology, Medicine and Surgery; the Osteoporosis Research Center, The Boys Town National Research Hospital and the Veteran’s Administration Hospital.

Neuroscience (Neuroanatomy, Neurobiology, Neurophysiology)
David He: Auditory neurobiology and neurophysiology
Tejbeer Kaur: Auditory neurobiology and neurophysiology
Garrett Soukup: Role of MicroRNAs in neurosensory systems
Peter Steyger: Auditory neurobiology and neurophysiology
Jian Zuo: Auditory neurobiology and neurophysiology

Hearing and Hearing loss
David He: Cochlear sensory hair cells, outer hair cell motility, age-related hearing loss
Ken Kramer: Role of cilia linkages to extracellular structures in inner ear function
Tejbeer Kaur: Innate immunity to hearing loss and neurodegeneration
Garrett Soukup: MicroRNAs in development and maintenance of sensory hair cells and neurons
Peter Steyger: Mechanisms of aminoglycoside antibiotics-induced hearing loss
Jian Zuo: Sensory hair cell protection and regeneration

Cancer Biology
Laura Hansen: Prevention and treatment of skin cancer
Brian North: Molecular mechanisms of cancer, aging and age-related diseases

Molecular Structure and Function of Peptides
Sandor Lovas: Peptide chemistry, Structural bioinformatics and proteomics, biophysical chemistry
David Smith: Chemical synthesis of peptides

Teaching Expertise in the Masters of Medical Sciences Program

Our faculty have an impressive range of expertise and students in this 11-month Master of Science degree program will have the opportunity to take courses in everything from human gross anatomy and neuroanatomy to pathology, surgery, radiology, and embryology. Our faculty work closely with our students, advising and mentoring throughout the course of study, as they share their collective passion of caring for the whole person.

Tyler E. Dunn, MS | Department of Medical Education

Kevin T. FitzGerald, SJ, PhD, PhD | Department of Medical Humanities

Leah D. Hunter, PhD | Department of Medical Education

Kenneth L. Kramer, PhD | Department of Biomedical Sciences

Carol S. Lomneth, PhD | Department of Medical Education

Venkatesh Govindarajan, PhD | Department of Medical Education

Cassandra Hays, PhD | Department of Medical Education

Anthony E. Kincaid, PT, PhD | Pharmacy Sciences

Leslie Kuhnel, DBe, MSHCE, MPA | Interdisciplinary Studies

Nicole Piemonte, PhD | Medical Humanities

Learn about the MMS program

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Masters in Biomedical Sciences

  • Carly Baker – affiliated with Dr. Shibata’s lab
  • Fred Millan – affiliated with Dr. Zuo’s lab
  • Caroline Pass – affiliated with Dr. Zuo’s lab
  • Mi Zhou - affiliated with Dr. Zuo’s lab


Masters in Medical Sciences

  • Michele Fujita
  • Christopher Grimont
  • Joseph Hrdlicka
  • April Mabie
  • Brooke Malchow
  • James Marchant
  • Kevin Nguyen
  • Kelsey Novak
  • Alexander Sjovold
  • Sydney Smith
  • Evan Sousa
  • Cameron Trout
  • Jessica Ung
  • Benjamin Volberding


PhD in Biomedical Sciences

  • Shreshtha Dash – affiliated with Dr. Steyger’s lab
  • Moynul Islam – Rotating labs
  • Rachel Johnson – affiliated with Dr. Hansen’s lab
  • Seraphine Kamayirese – affiliated with Dr. Lovas’ lab
  • Rotachi Nlekwuwa affiliated with Dr. North’s lab
  • Renju Pun – affiliated with Dr. North’s lab
  • Samadhi Warnakulasooriya – affiliated with Dr. North’s lab


MD/MS/PhD Students

  • David Doss – affiliated with Dr. Mahapatra’s lab
  • Margaret McCann – affiliated with Dr. Zuo’s lab
  • Molly Kubesh – affiliated with Dr. Zuo’s lab
  • Jonathan Fleegel – affiliated with Dr. Zuo’s lab
  • Justin Rudd – affiliated with Dr. Hansen’s lab

Ready to start your next steps?